High-density mapping of the MHC identifies a shared role for HLA-DRB1*01:03 in inflammatory bowel diseases and heterozygous advantage in ulcerative colitis.

Goyette, P, Boucher G, Mallon D, Ellinghaus E, Jostins L, Huang H, […], Franke A, Rioux J D (2015); Nature Genetics, 47:172-179. doi: 10.1038/ng.3176